Jennifer Neuwald Associate Professor

Office: Biology 342

Phone: (970) 491-2796

Google Scholar:


  • Ph.D., Washington University in St. Louis
  • M.Sc., San Diego State University
  • B.Sc., Michigan State University



  • PhD, Washington University in St. Louis
  • MSc, San Diego State University
  • BSc, Michigan State University


I am an evolutionary ecologist interested in using a multidisciplinary approach to investigate how environmental variation and evolutionary processes converge to influence the patterns of demographic, genetic, and genomic variation in natural populations, especially those of conservation concern. Projects that I am currently engaged in include:

  • effects of forest fires on the demography and genetic signatures of collared lizards
  • repeated patterns of evolution in founding populations of collared lizards
  • influence of metapopulation dynamics on estimates of variance effective size of populations
  • patterns in organisms' abilities to acclimate to environmental variance


  • LIFE102 (Attributes of Living System)
  • BZ220 (Evolution)
  • BZ329 (Herpetology) - variable
  • FW558 (Conservation Genetics of Wild Vertebrates) - CSU On-line
  • BZ495, BZ498, BZ487 (non-regular courses)  

CSU Leadership, Initiatives, and Pedagody:

Select Publications:

  • Havrid, J., Neuwald, J., Shah, A., Mauro, A., Marshall, C., Ghalambor, C. (2020) Distinguishing between active plasticity due to thermal acclimation and passive plasticity due to Q10 effects: Why methodology matters. Functional Ecology. 34(5), 1015-1028.
  • Valenzuela, N., Literman, R., Neuwald, J., Mizoguchi, B., Iverson, J., Riley, J., Litzgus, J. (2019) Extreme thermal fluctuations from climate change to unexpectedly accelerate demographic collapse of vertebrates with temperature-dependent sex determination. Scientific Reports. 9(1), 4254.
  • Radhakrishnan, S., Literman, R., Neuwald, J. L., & Valenzuela, N. (2018) Thermal Response of Epigenetic Genes Informs Turtle Sex Determination with and without Sex Chromosomes. Sexual Development. 12(6), 308.
  • Piaggio, A.J., Osorio, I.A., Ramirez, A.J., Neuwald, J.L., et al. (2017) Genetic demography at the leading-edge of the distribution of a potentially invasive rabies virus vector. Ecology and Evolution, 7(14): 5343.
  • Radhakrishnan, S., Valenzuela, N., Literman, R., Neuwald, J., et al. (2017) Transcriptomic responses to environmental temperature by turtles with thermosensitive and genotypic sex determination assessed by RNAseq inform the genetic architecture of embryonic gonadal development. PLoS ONE, 12(3): e0172044.
  • Neuwald, J.L. and A.R. Templeton (2013) Genetic restoration in the eastern collared lizard under prescribed woodland burning. Molecular Ecology, 22:3666-3679.
  • Valenzuela N., J.L. Neuwald, R. Literman (2012) Transcriptional evolution underlying vertebrate sexual development. Developmental Dynamics. DOI: 10.1002/dvdy.23897.
  • Neuwald, J.L. and N. Valenzuela (2011) The lesser known challenge of climate change: Thermal variance and sex-reversal in vertebrates with temperature-dependent sex determination. PLoS One, 6(3): e18117. doi:10.1371/journal.pone.0018117.
  • Templeton, A.R., H. Brazeal, and J.L. Neuwald (2011) The transition from isolated patches to a metapopulation in the eastern collared lizard in response to prescribed woodland fires. Ecology, 92:1736-1747.
  • Neuwald, J.L. (2010) Population isolation exacerbates conservation genetic concerns in the endangered Amargosa vole, Microtus californicus scirpensis. Biological Conservation. 143: 2028-2038.
  • Conroy, C.J. and J.L. Neuwald (2008) Phylogeographic study of the California vole, Microtus californicus. Journal of Mammalogy. 89:755-767.
  • Templeton, A.R., J.L. Neuwald, et al. (2007) Invited Mini-Review: Restoring demographic processes in translocated populations of collared lizards in the Missouri Ozarks through prescribed forest fires. Israel Journal of Ecology and Evolution. 53:179-196.