The wood frog is one of the most widely distributed species in North America. Within its range, it is able to exploit a wider range of conditions than other species. Part of their success stems from unusual tolerance to closed canopy conditions within their breeding environments. In spite of cool temperatures and low productivity, larvae are able to perform More remarkably, wood frogs undergo micro geographic scale divergence in a variety of traits. These shifts counteract environmental gradients that can impair performance and imply either that wood frogs can evolve rapidly to deal with changing conditions within breeding ponds or that animals may focus migrations to match phenotype to habitat. Under either scenario, the emerging understanding of niche breadth in this species suggests strong, real time interaction between ecological and evolutionary response to environmental heterogeneity.